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Most di-thiol Grx use monothiol mechanisms that take place in two steps: In the first step, the nucleophilic thiolate anion attacks the S -glutathionylated substrate protein, resulting in reduction of the mixed disulfide and the S -glutathionylated Grx (Grx-SSG) intermediate

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One prominent category of natural extracts includes flavonoids, a diverse group of polyphenolic compounds found in fruits, vegetables, and beverages like tea