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Introduction Reactive oxygen species (ROS), once perceived as toxic by-products, were known to cause oxidative damage in cells (Mittler et al., 2 O 2 is an endogenous ROS species known to play a dual role in plants, where it is beneficial at low concentrations but lethal at higher levels (Petrov and Van Breusegem, 2 O 2 acts as signaling molecule inducing the signal transduction mechanism to produce various cellular responses