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Because the oxidationreduction of Fe 2+ to Fe 3+ requires electron transfer, iron is involved in the electron migration between substrates ( 2+ or Fe 3+ ions and hydrogen peroxide (H 2 O 2 ) to produce highly reactive hydroxyl radicals (Fenton, 1894), which increased ROS level in cells and triggered the peroxidation of polyunsaturated fatty acids (PUFAs) or saturated fatty acids in membrane lipids (Lachaier et al., 2014)

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