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Together, these results suggested to us the possibility that NAC-induced roGFP2 oxidation is caused by sulfane sulfur (S 0 ) species created inside mitochondria, based on the following consideration (summarized in Figure 1E): It is known that H 2 S is very efficiently oxidized by sulfide:quinone oxidoreductase (SQR) to yield hydropersulfides inside the mitochondrial matrix (Libiad et al., 2014)

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Intranasal GSH (Phase IIb): Randomized, double-blind, placebo-controlled study (n=45) tested intranasal reduced GSH 100 or 200 mg three times daily for 12 weeks

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